LONDON - The Jaguars will be without starting tight end Julius Thomas on Sunday.

Thomas has been battling an elbow injury this week and wasn't a full participant in practice at any point during preparation for the Colts.

Joining Thomas on the sidelines will be starting Otto linebacker Dan Skuta, who is dealing with a hip injury.

Defensive end Jared Odrick (tricep) and backup tight end Ben Koyack will also miss the game. Both Koyack and Odrick stayed in Jacksonville and did not travel to London.

Veteran defensive lineman Tyson Alualu will start in place of Odrick at defensive end. Rookie linebacker Myles Jack will start at Otto for Skuta, despite only playing in small doses on defense in the first three games.

Defensive end Chris Smith is active for the first time this season.

Here is the entire inactive list for the Jaguars in Week 4:

- CB Dwayne Gratz

- TE Ben Koyack

- DE Jared Odrick

- TE Julius Thomas

- LB Dan Skuta

- OG Chris Reed

- QB Brandon Allen

Follow Mike Kaye on Twitter at @Mike_E_Kaye.